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postfix delivers mail locally via LMTP. I believe the standard LMTP port is 7025.
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Yes! That's the magic number! Thank you. I opened that port and it's working fine now.
It was rather frustrating that the error message wasn't giving me a port number. If it's not mentioned in the multi-server install guide already (I couldn't find it), it should be mentioned that the Mailbox server must have port 7025 open to receive email. I would suspect that anyone installing a multi-server configuration on a secured network would have the same issue.